Systematic problems surrounding Howarthia melli (Forster) (Lepidoptera: Lycaenidae: Theclinae), with description of a new species and a review of Rhododendron-association in Lycaenid butterflies

Yu Feng Hsu, Dongsun Ding, Shen Horn Yen, Zu Qi Qian

Research output: Contribution to journalArticle

3 Citations (Scopus)


Within the genus Howarthia, Howarthia melli (Forster) is the most frequently reported species across the southern provinces of China. An examination of the type series of H. melli, in combination with results of rearing experiments, reveals that three species have been frequently confused under the name H. melli. The three are superficially similar but not closely related to each other within the genus. H. melli is much more restricted in distribution than previously thought, with most literature records referring to the other two species, Howarthia cheni Chou & Wang and an undescribed species, Howarthia cyanea sp. n., described herein. We provide a discussion of the relationships of these species and a review of Rhododendron-association in Lycaenidae.

Original languageEnglish
Pages (from-to)651-666
Number of pages16
JournalAnnals of the Entomological Society of America
Issue number4
Publication statusPublished - 2004 Jul 1



  • China
  • Howarthia cheni
  • Howarthis cyanea
  • New species
  • Theclini

ASJC Scopus subject areas

  • Insect Science

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