TY - JOUR
T1 - Revision of Pazala Moore, 1888
T2 - The Graphium (Pazala) mandarinus (Oberthür, 1879) Group, with Treatments of Known Taxa and Descriptions of New Species and New Subspecies (Lepidoptera: Papilionidae)
AU - Hu, Shao Ji
AU - Cotton, Adam M.
AU - Condamine, Fabien L.
AU - Duan, Kuang
AU - Wang, Rong Jiang
AU - Hsu, Yu Feng
AU - Zhang, Xin
AU - Cao, Jun
N1 - Funding Information:
The authors wish to thank the following persons who kindly allowed us to access public and private collections, photographed or provided specimens and helped collect samples for DNA analysis (in alphabetic order of surnames): Peter Andrews (UK), Danny Burk (South Bend, Indiana, USA), David Cassat (Australia), Jia-Lin Chen (Guangzhou, China), Zhi-Bing Chen (Shanghai, China), Hideyuki Chiba (Japan), Christoph Häuser (Berlin, Germany), James Hogan (Oxford University Museum of Natural History, UK), Ping Hu (Peking University, Beijing, China), Blanca Huertas (BMNH, London, UK), Zhuo-Heng Jiang (Kunming, China), Toshihiko Katayama (Tochigi, Japan), Gerardo Lamas (Museo de Historia Natural, Lima, Peru), Song-Yun Lang (Chongqing, China), Yu-Fei Li (Xi’an, China), Guang Liu (Guangzhou, China), Philip Yik-Fui Lo (Kadoorie Farm and Botanic Garden, Hong Kong, China), Wei-Wei Mao (Shanghai, China), Geoff Martin (BMNH, London, UK), Jan Moonen (Cadier en Keer, The Netherlands), Yasusuke Nishiyama (Tokyo, Japan), Tommaso Racheli (Rome, Italy), Peter Smetacek (Nainital, India), Campbell Smith (BMNH, London, UK), Axel Steiner (Stuttgart, Germany), Dieter Stüning (ZFMK, Bonn, Germany), Prasobsuk Sukkit (†) (Chiang Mai, Thailand), Trustees of the Natural History Museum (London, UK), R. I. Vane-Wright (BMNH, London, UK), Bob Worthy (BMNH, London, UK), Chuan-Hui Yi (Southwest Forestry University, Kunming, China), Li-Kun Zhang (Kunming Institute of Zoology, CAS, Kunming, China), and Jian-Qing Zhu (Shanghai, China). Special thanks must be given to Rui-Xuan Chen-Liu (Fuzhou University, Fuzhou, China) and Hui-Hong Zhang (Yunnan University, Kunming, China) for discovering the host plant of Graphium (Pazala) confucius sp. nov. and bringing back valuable material, and to Laohegou Nature Reserve and Tangjiahe Nature Reserve in N. Sichuan for assistance in collecting G. (P.) sichuanica. This study was funded by the National Key R&D Programme of China (2017YFC0505200), the NSFC Programme of China (41761011) and the Biodiversity Conservation Programme of the Ministry of Ecology and Environment, China (China-BON Butterflies) (SDZXWJZ01013).
Publisher Copyright:
Copyright © 2018 Magnolia Press.
PY - 2018/6/28
Y1 - 2018/6/28
N2 - The previously recognised closely related species Graphium (Pazala) mandarinus (Oberthür, 1879) and G. (P.) sichuanica (Koiwaya, 1993) are shown to comprise seven species as a result of both molecular and morphological analysis. Molecular dating analysis is also performed on the mandarinus group in order to investigate the divergence time of the taxa. Two taxa, G. (P.) garhwalica (Katayama, 1988) stat. nov. and G. (P.) paphus (de Nicéville, 1886) stat. nov., are raised from subspecific to specific status; G. (P.) hoeneanus Cotton & Hu nom. nov., stat. rev. is separated from sichuanica at species level; and two previously unrecognised new species, G. (P.) daiyuanae Hu, Zhang & Cotton sp. nov. and G. (P.) confucius Hu, Duan & Cotton sp. nov. are described from Vietnam and China respectively, the latter being sympatric with nominate G. (P.) mandarinus. The identity of the lectotype of G. (P.) mandarinus is confirmed and a lectotype is designated for the taxon Papilio Glycerion Gray, 1831. A new subspecies of G. (P.) mandarinus is described from western Yunnan and northern Myanmar, G. (P.) mandarinus stilwelli Cotton & Hu ssp. nov.
AB - The previously recognised closely related species Graphium (Pazala) mandarinus (Oberthür, 1879) and G. (P.) sichuanica (Koiwaya, 1993) are shown to comprise seven species as a result of both molecular and morphological analysis. Molecular dating analysis is also performed on the mandarinus group in order to investigate the divergence time of the taxa. Two taxa, G. (P.) garhwalica (Katayama, 1988) stat. nov. and G. (P.) paphus (de Nicéville, 1886) stat. nov., are raised from subspecific to specific status; G. (P.) hoeneanus Cotton & Hu nom. nov., stat. rev. is separated from sichuanica at species level; and two previously unrecognised new species, G. (P.) daiyuanae Hu, Zhang & Cotton sp. nov. and G. (P.) confucius Hu, Duan & Cotton sp. nov. are described from Vietnam and China respectively, the latter being sympatric with nominate G. (P.) mandarinus. The identity of the lectotype of G. (P.) mandarinus is confirmed and a lectotype is designated for the taxon Papilio Glycerion Gray, 1831. A new subspecies of G. (P.) mandarinus is described from western Yunnan and northern Myanmar, G. (P.) mandarinus stilwelli Cotton & Hu ssp. nov.
KW - China
KW - India
KW - Leptocircini
KW - New species
KW - Pazala
KW - Vietnam
UR - http://www.scopus.com/inward/record.url?scp=85049240763&partnerID=8YFLogxK
UR - http://www.scopus.com/inward/citedby.url?scp=85049240763&partnerID=8YFLogxK
U2 - 10.11646/zootaxa.4441.3.1
DO - 10.11646/zootaxa.4441.3.1
M3 - Review article
C2 - 30313994
AN - SCOPUS:85049240763
SN - 1175-5326
VL - 4441
SP - 401
EP - 446
JO - Zootaxa
JF - Zootaxa
IS - 3
ER -